On
"Subspecies" of the Canada Goose
by Gerry Rising
(This is an article submitted to The
Kingbird in 2003.
It is a revised version of an article that appeared in The Prothonotary 68 (12):131-132.)
I
am concerned about the interest being expressed by New York State observers in
the smaller Canada Geese occurring here. These are, according to them, Branta
canadensis hutchinsii, a subspecies
termed the Hutchins Goose in Bent's Life Histories of North American
Waterfowl. (Elsewhere it is named
Hutchins', Hutchins's or even Richardson's Goose.) I certainly do not argue
with this identification, but I wince each time I see these birds reported. In
this article I explain why.
But
first an aside: I have no problem with our differentiating the Horned Lark and
Palm Warbler subspecies or the various hybrid forms of the Golden-winged and
Blue-winged Warblers. There are, I believe, good arguments for these
identifications. However, with game birds and in particular Canada Geese, I
urge caution in raising this issue of races.
Now
to my point: Surely most of us would agree that we have today a few too many
Canada Geese soiling our ball fields and corporate lawns, ruining our golf
course greens and driving other species out of our marshes. What we should
understand is that interest in the various forms of Canada Geese contributes to
this overpopulation.
How
many birders realize that goose-hunting seasons have been severely curtailed
because of concern for another so-called race of Canada Goose, B. c. maxima, a subspecies that is not even recognized by
taxonomists of the American Ornithologists Union?
These
larger geese were popularized by Harold C. Hansen's The Giant Canada Goose (Carbondale, IL: Southern Illinois University Press,
1965, rev. 1997). In this book, which has been termed by one wildlife manager
"one of the finest waterfowl monographs ever written," Hansen makes
the case for protecting these big geese ‹ hunters' favorites for obvious
reasons ‹ through season restrictions: "Adjustment of the opening and
closing dates of the hunting season," he says, "is one of the
principal means of regulating the size and selectivity of waterfowl kills. It
should, therefore,...be pointed out that because the fall migration movements
of the giant Canadas are fully a month behind the migration of the race interior, late hunting seasons will tend to jeopardize stocks
of maxima...."
And
so, at least partly influenced by Hansen, goose seasons have been reduced. As
my Ohio colleague, Bill Whan, says: "It is an article of belief among
wildlife officials that their precious maximae 'migrate' a month later than other forms. Thus here
in Ohio we have an 'early goose season' in September (coinciding with the teal
season pretty much), which is ballyhooed as a way to control nuisance flocks of
resident geese without imperiling any of the truly wild and migratory Canadas,
which allegedly come through beginning in mid-October." But even this goes
wrong. Bill continues: "This year the truly migrant race interior - with orange collars - (universally recognized as
genuinely in trouble) came through a bit 'earlier' than usual...to be greeted
by a hail of shot."
Hanson
fought hard to have the American Ornithologists Union recognize his beloved
giants, but Alexander Wetmore and his taxonomic colleagues would have no part
of his arguments. The only mention of the maxima subspecies in any of the seven A.O.U. Checklists is in the 5th (1957). There it is listed as probably extinct. More important,
especially as regards our local Canadas is the 6th (1983) Checklist. In it no subspecies is listed! And this note
appears: "The northern populations of small Canada Geese have been
variously treated taxonomically as three separate species, B. hutchinsii (Richardson, 1832) [Hutchins'
or Richardson's Goose, 172.3], B. minima Ridgway, 1885 [Cackling
Goose, 172.2], and B. leucopareia (Brandt, 1836) [Tundra Goose,
172.1]; as a single species under the name B. hutchinsii [Cackling
Goose]; or as one or more subspecies of B. Canadensis. Consideration of the entire complex as a single
species seems best for the present." (The underscoring of these
important points is mine.) To bring this up to date: The 7th Checklist (1998) follows the 6th in listing no subspecies. Clearly all of these size
differences are currently considered a cline.
Why,
despite this, Kevin Griffith mentions B. c. maxima in the 1998 Bull's Birds of New York State escapes me. Consider, however, what he says:
"Resident breeding geese probably include some of both [the canadensis and interior subspecies] and a large proportion of maxima, or 'Giant' Canada Goose." What a turn-around:
from Hansen's endangered race needing protection to the lawn pests of Griffith
and, as it happens, of current waterfowl managers! (pers. comm., Paul Hess) Has no one thought of the possibility
that these 'giant' geese are bigger and fatter simply because they are lazy,
pampered and overfed?
My
point here is that attention to game bird subspecies gives wildlife managers an
excuse to tamper with what I consider an already far too short goose season.
While they concern themselves with niggling subpopulations, we're being
inundated by the species as a whole. And we certainly don't want to provide an
excuse for a season further curtailed to address the smaller numbers of hutchinsii.
An
indication of just how far this race issue may be taken is found in the revised
(1997) edition of Hansen's book. In it he insists on page xvii that "As a
result of over 30 years of research on the taxonomy and biogeography of the
white-cheeked geese, at least 186 geographic races (subspecies) of these geese
will be recognized in a yet-to-be published work." He comments on this
work in progress in a footnote: "The White-cheeked geese (Branta
canadensis, B. maxima, B. hutchinsii, B. leucopareia, and B. minima): Taxonomy, Ecophysiographic Relationships,
Biogeography, and Evolutionary Considerations. (Yet to be published.) Late in
the preparation of this work on the white-cheeked geese, it was realized that
the giant Canada goose (Branta canadensis maxima) was in reality a separate species -- Branta
maxima -- and that it should be
given a more appropriate common name -- the giant prairie goose. Seven
geographic races or subspecies will be described."
He
adds to this in Appendix I on pages 207-208: "The 83 races at least of
Canada geese (Branta canadensis)
that will be recognized by the author (Hanson, yet to be published) represent a
series of staircase clines in respect to size, proportions, color, and color
patterns -- that is, each racial population exhibits a discrete combination of
these characters that are related to the latitude, climate, and ecology of
their isolated breeding grounds. Stated another way, the mosaic of characters
associated with the racial populations of white-cheeked geese are not accompanied
by 'intergrades' or individuals that represent a blending of the characters of
the nearest racial populations."
It
is easy to be confused by all that nonsense. What he appears to be saying is
that he is separating out from the Branta genus (because Brant, Branta bernicla, is no longer included in its own genus) these
so-called white-cheeked geese for which he identifies "at least 186"
races, 83 of which are races grouped under the species Branta canadensis. This leaves us with over 100 races that would have
to be identified with new species, one of which he identifies as Branta
maxima, no longer the Giant Canada
Goose, but now the Giant Prairie Goose. And his "new species" has
only seven of those 100 races, thus suggesting still more full species. Hansen
certainly deserves some kind of prize (offered by listers?) as the champion
among all splitters.
Of
course this all flies in the face of John W. Aldrich's 1946 article
"Speciation in the White-cheeked Geese" (Wilson Bulletin 58(2): 94-103) which identifies, among 254 museum
specimens examined, only seven races, none of them maxima; and the most subspecies ever considered by an
A.O.U. Checklist: ten in the 5th
(1957).
I
join Bill Whan in wondering if "all those goose-droppings in the water
have had some neurotoxic effects" on a few game managers. Whether or not
that is the case, I counsel extreme caution in calling attention to nonexistent
subspecies to avoid even the possibility of contributing further to our
increasingly serious Canada Goose overpopulation problem.
A
final note: Despite all I have said here, it is, I believe, quite reasonable to
differentiate between our tame, non-migratory geese and the wild geese that
migrate through this country and Canada. What we really need is a national
campaign to rid ourselves of the pests that we have created in the same way
that waterfowl managers are finally seeking to control cormorants and Mute
Swans ‹ by activities like egg-oiling and removal (a.k.a. killing). Then goose
seasons could be scheduled in response to wild populations. Sadly, however,
even the consideration of such actions divides our community.